Abstract
Physiologists have made few studies on hummingbirds despite the attention that
these birds deserve because of their small size and their ability to fly in a small con-
tainer. By measuring oxygen consumption I have been able to calculate their rate of
metabolism at rest and in flight and their energy needs over a 24-hour period. It has
also been possible to measure the depth of the torpidity that some investigators have
suspected occurs and to make a rough estimate of the non-stop flight range of hum-
mingbirds.
METHODS
Five Anna (Calypte anna) and two Allen (Selasphorus sasin sas
were studied. These were captured as nestlings or as adults and wer
ture of water, sugar, milk, and Mellon's food, or a mixture of wat
strained liver. Rate of metabolism was measured by oxygen con
circuit volumetric apparatus similar to that described by Morrison
minations during which no flying was wanted, the animal chamber
5 inches in diameter and 3 inches high (inside measurements). For d
ing which the bird was expected to fly, a bell jar 6 inches in diamete
was used. Suitable surfaces of soda lime and calcium chloride we
and the large bell jar was equipped with a fan to stir up the air and
tion of carbon dioxide and water vapor. These chambers were conne
of oxygen and when in use were submerged in a water bath to keep
inside them constant. Food was provided for the birds in the bell j
special cases. When a bird was confined in the apparatus for se
weighed both before and after, and the weight used for calculation
at any period during the experiment was arrived at by interpolation
ments the animal was weighed only at the end. The oxygen consum
this paper are expressed in cubic centimeters of gas at 0OC. and 76
Acknowledgment.-I am much indebted to Mr. and Mrs. Eric
plying me with the Allen Hummingbirds used and for much valuab
for hummingbirds in captivity.
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